The Palaeo-Agulhas Plain formed an important habitat exploited by Pleistocene hunter-gatherer populations during periods of lower sea level. This productive, grassy habitat would have supported numerous large-bodied ungulates accessible to a population of skilled hunters with the right hunting technology. It also provided a potentially rich location for plant food collection, and along its shores a coastline that moved with the rise and fall of sea levels. The rich archaeological and paleontological records of Pleistocene sites along the modern Cape south coast of South Africa, which would have overlooked the Palaeo-Agulhas Plain during Pleistocene times of lower sea level, provides a paleoarchive of this extinct ecosystem. In this paper, we present a first order illustration of the “palaeoscape modeling” approach advocated by Marean et al. (2015). We use a resourcescape model created from modern studies of habitat productivity without the Palaeo-Agulhas Plain. This is equivalent to predominant Holocene conditions. We then run an agent-based model of the human foraging system to investigate several research questions. Our agent-based approach uses the theoretical framework of optimal foraging theory to model human foraging decisions designed to optimize the net caloric gains within a complex landscape of spatially and temporally variable resources. We find that during the high sea-levels of MIS 5e (+5-6 m asl) and the Holocene, the absence of the Plain left a relatively poor food base supporting a much smaller population relying heavily on edible plant resources from the current Cape flora. Despite high species diversity of plants with edible storage organs, and marine invertebrates, encounter rates with highly profitable resources were low. We demonstrate that without the Palaeo-Agulhas Plain, human populations must have been small and low density, and exploited plant, mammal, and marine resources with relatively low caloric returns. The exposure and contraction of the Palaeo-Agulhas Plain was likely the single biggest driver of behavioral change during periods of climate change through the Pleistocene and into the transition to the Holocene.

This model is an extension of the Artificial Long House Valley (ALHV) model developed by the authors (Swedlund et al. 2016; Warren and Sattenspiel 2020). The ALHV model simulates the population dynamics of individuals within the Long House Valley of Arizona from AD 800 to 1350. Individuals are aggregated into households that participate in annual agricultural and demographic cycles. The present version of the model incorporates features of the ALHV model including realistic age-specific fertility and mortality and, in addition, it adds the Black Mesa environment and population, as well as additional methods to allow migration between the two regions.

As is the case for previous versions of the ALHV model as well as the Artificial Anasazi (AA) model from which the ALHV model was derived (Axtell et al. 2002; Janssen 2009), this version makes use of detailed archaeological and paleoenvironmental data from the Long House Valley and the adjacent areas in Arizona. It also uses the same methods as the original AA model to estimate annual maize productivity of various agricultural zones within the Long House Valley. A new environment and associated methods have been developed for Black Mesa. Productivity estimates from both regions are used to determine suitable locations for households and farms during each year of the simulation.