A model to investigate the Evolution of Conditional Cooperation in a Spatial Public Goods Game. We consider two conditional cooperation strategies: one based on thresholds (Battu & Srinivasan, 2020) and another based on independent decisions for each number of cooperating neighbors. We examine the effects of productivity and conditional cooperation criteria on the trajectory of cooperation. Cooperation is evolving with no need for additional mechanisms apart from spatial structure when agents follow conditional strategies. We confirm the positive influence of productivity and cluster formation on the evolution of cooperation in spatial models. Results are robust for the two types of conditional cooperation strategies.

The emergence of cooperation in human societies is often linked to environmental constraints, yet the specific conditions that promote cooperative behavior remain an open question. This study examines how resource unpredictability and spatial dispersion influence the evolution of cooperation using an agent-based model (ABM). Our simulations test the effects of rainfall variability and resource distribution on the survival of cooperative and non-cooperative strategies. The results show that cooperation is most likely to emerge when resources are patchy, widely spaced, and rainfall is unpredictable. In these environments, non-cooperators rapidly deplete local resources and face high mortality when forced to migrate between distant patches. In contrast, cooperators—who store and share resources—can better endure extended droughts and irregular resource availability. While rainfall stochasticity alone does not directly select for cooperation, its interaction with resource patchiness and spatial constraints creates conditions where cooperative strategies provide a survival advantage. These findings offer broader insights into how environmental uncertainty shapes social organization in resource-limited settings. By integrating ecological constraints into computational modeling, this study contributes to a deeper understanding of the conditions that drive cooperation across diverse human and animal systems.

NeoCOOP is an iteration-based ABM that uses Reinforcement Learning and Artificial Evolution as adaptive-mechanisms to simulate the emergence of resource trading beliefs among Neolithic-inspired households.

The model explores food distribution patterns that emerge in a small-scale non-agricultural group when individuals follow a set of spatially explicit sharing interaction rules derived from a theory on the evolution of the egalitarian social instinct.

Cooperation is essential for all domains of life. Yet, ironically, it is intrinsically vulnerable to exploitation by cheats. Hence, an explanatory necessity spurs many evolutionary biologists to search for mechanisms that could support cooperation. In general, cooperation can emerge and be maintained when cooperators are sufficiently interacting with themselves. This communication provides a kind of assortment and reciprocity. The most crucial and common mechanisms to achieve that task are kin selection, spatial structure, and enforcement (punishment). Here, we used agent-based simulation models to investigate these pivotal mechanisms against conditional defector strategies. We concluded that the latter could easily violate the former and take over the population. This surprising outcome may urge us to rethink the evolution of cooperation, as it illustrates that maintaining cooperation may be more difficult than previously thought. Moreover, empirical applications may support these theoretical findings, such as invading the cooperator population of pathogens by genetically engineered conditional defectors, which could be a potential therapy for many incurable diseases.

The model explores food distribution patterns that emerge in a small-scale non-agricultural group when sharing individuals engage in intentional consumption leveling with a given probability.

This model replicates the Axelrod prisoner’s dilemma tournaments. The model takes as input a file of strategies and pits them against each other to see who achieves the best payoff in the end. Change the payoff structure to see how it changes the tournament outcome!

Previous work with the spatial iterated prisoner’s dilemma has shown that “walk away” cooperators are able to outcompete defectors as well as cooperators that do not respond to defection, but it remains to be seen just how robust the so-called walk away strategy is to ecologically important variables such as population density, error, and offspring dispersal. Our simulation experiments identify socio-ecological conditions in which natural selection favors strategies that emphasize forgiveness over flight in the spatial iterated prisoner’s dilemma. Our interesting results are best explained by considering how population density, error, and offspring dispersal affect the opportunity cost associated with walking away from an error-prone partner.

This study investigates a possible nexus between inter-group competition and intra-group cooperation, which may be called “tribalism.” Building upon previous studies demonstrating a relationship between the environment and social relations, the present research incorporates a social-ecological model as a mediating factor connecting both individuals and communities to the environment. Cyclical and non-cyclical fluctuation in a simple, two-resource ecology drive agents to adopt either “go-it-alone” or group-based survival strategies via evolutionary selection. Novelly, this simulation employs a multilevel selection model allowing group-level dynamics to exert downward selective pressures on individuals’ propensity to cooperate within groups. Results suggest that cooperation and inter-group conflict are co-evolved in a triadic relationship with the environment. Resource scarcity increases inter-group competition, especially when resources are clustered as opposed to widely distributed. Moreover, the tactical advantage of cooperation in the securing of clustered resources enhanced selective pressure on cooperation, even if that implies increased individual mortality for the most altruistic warriors. Troubling, these results suggest that extreme weather, possibly as a result of climate change, could exacerbate conflict in sensitive, weather-dependent social-ecologies—especially places like the Horn of Africa where ecologically sensitive economic modalities overlap with high-levels of diversity and the wide-availability of small arms. As well, global development and foreign aid strategists should consider how plans may increase the value of particular locations where community resources are built or aid is distributed, potentially instigating tribal conflict. In sum, these factors, interacting with pre-existing social dynamics dynamics, may heighten inter-ethnic or tribal conflict in pluralistic but otherwise peaceful communities.

For special issue submission in JASSS.

Righi S., Takacs K., Social Closure and the Evolution of Cooperation via Indirect Reciprocity, Resubmitted after Revisions to Scientific Reports