The Palaeo-Agulhas Plain formed an important habitat exploited by Pleistocene hunter-gatherer populations during periods of lower sea level. This productive, grassy habitat would have supported numerous large-bodied ungulates accessible to a population of skilled hunters with the right hunting technology. It also provided a potentially rich location for plant food collection, and along its shores a coastline that moved with the rise and fall of sea levels. The rich archaeological and paleontological records of Pleistocene sites along the modern Cape south coast of South Africa, which would have overlooked the Palaeo-Agulhas Plain during Pleistocene times of lower sea level, provides a paleoarchive of this extinct ecosystem. In this paper, we present a first order illustration of the “palaeoscape modeling” approach advocated by Marean et al. (2015). We use a resourcescape model created from modern studies of habitat productivity without the Palaeo-Agulhas Plain. This is equivalent to predominant Holocene conditions. We then run an agent-based model of the human foraging system to investigate several research questions. Our agent-based approach uses the theoretical framework of optimal foraging theory to model human foraging decisions designed to optimize the net caloric gains within a complex landscape of spatially and temporally variable resources. We find that during the high sea-levels of MIS 5e (+5-6 m asl) and the Holocene, the absence of the Plain left a relatively poor food base supporting a much smaller population relying heavily on edible plant resources from the current Cape flora. Despite high species diversity of plants with edible storage organs, and marine invertebrates, encounter rates with highly profitable resources were low. We demonstrate that without the Palaeo-Agulhas Plain, human populations must have been small and low density, and exploited plant, mammal, and marine resources with relatively low caloric returns. The exposure and contraction of the Palaeo-Agulhas Plain was likely the single biggest driver of behavioral change during periods of climate change through the Pleistocene and into the transition to the Holocene.

This model is an application of Brantingham’s neutral model to a real landscape with real locations of potential sources. The sources are represented as their sizes during current conditions, and from marine geophysics surveys, and the agent starts at a random location in Mossel Bay Region (MBR) surrounding the Archaeological Pinnacle Point (PP) locality, Western Cape, South Africa. The agent moves at random on the landscape, picks up and discards raw materials based only upon space in toolkit and probability of discard. If the agent happens to encounter the PP locality while moving at random the agent may discard raw materials at it based on the discard probability.

This model allows for the investigation of the effect spatial clustering of raw material sources has on the outcome of the neutral model of stone raw material procurement by Brantingham (2003).

This model simulates a foraging system based on Middle Stone Age plant and shellfish foraging in South Africa.

The ForagerNet3_Demography model is a non-spatial ABM designed to serve as a platform for exploring several aspects of hunter-gatherer demography.

This model represents technological and ecological behaviors of mobile hunter-gatherers, in a variable environment, as they produce, use, and discard chipped stone artifacts. The results can be analyzed and compared with archaeological sites.

ForagerNet3_Demography_V2 is a non-spatial ABM for exploring hunter-gatherer demography. This version (developed from FN3D_V1) contains code for calculating the ratio of old to young adults (the “OY ratio”) in the living and dead populations.

ForagerNet3_Demography is a non-spatial ABM for exploring hunter-gatherer demography. Key methods represent birth, death, and marriage. The dependency ratio is an imporant variable in many economic decisions embedded in the methods.